Journal of Evolutionary Biology 2001, 14 (<), .

Supplementary material for pages 219-227

Appendix A

Details on the placement of additional genera


GenusEvidence for placementDecision
Afrosyrphusrelated to Epistrophe (Vockeroth, 1969)sister-group to Epistrophe
AlipumilioPosition supported by larval characters (Rotheray, unpubl. data). in Eumerus clade
Following Thompson (1972, p.125), we place it with Nausigaster 
(but see below) as sister-group to Eumerus. In Eumerus group 
(Hippa, 1978, p.18). 
Allobacchasubgenus of Baccha (Knutson et al., 1975). In the Syrphini sister-group to Baccha
(Thompson & Vockeroth, 1989) 
Ameroxylotasister-group to Brachypalpus (Hippa, 1978)sister-group to Brachypalpus
Aneriophorarelated to Temnostoma (Thompson, 1972, p.163). Usually placed with Macrozelima and Teuchocnemis
considered related to Criorhina group, but separated from them by a 
series of characters; resembles Teuchocnemis (Hippa, 1978, p.11) 
Arctosyrphusclosely related to Anasimyia group, since placed as a subgenus in Anasimyia group
there (Thompson, 1989) 
Argentinomyiarelated to Microdon (Thompson et al., 1976)in Microdon group
Asemosyrphusrelated to Helophilus (Thompson et al., 1976). in Anasimyia group
Put as subgenus of Anasimyia by Thompson (1989) 
Asiodideasister-group to Dideomima, and in the Didea group in Didea group
(Vockeroth, 1969, p.106) 
Austalisrelated to Eristalis (Thompson & Vockeroth, 1989)in Eristalis group
Austroasciarelated to Sphegina-Neoascia (Thompson, 1989)sister-group of Sphegina
Austrocheilosiarelated to Eumerus (Thompson & Vockeroth, 1989)in Eumerus group
Austrophilusrelated to Helophilus (Thompson & Vockeroth, 1989)in Helophilus group
Axonarelated to Eristalis (Thompson & Vockeroth, 1989)in Eristalis group
Azypetiasister-group to Merodon, according to Thompson (1972, p.125). sister-group to Merodon
In Eumerus group (Hippa, 1978, p.18) 
Cacoceriaaberrant xylotine (Thompson, 1972, p.180), perhaps sister-group to placed in the polytomy close to Brachyopa
Temnostoma (p 196). In the chrysogasterines 
(Thompson et al., 1976). Related to Brachyopa and the 
chrysogasterines (Thompson, 1989) 
Chamaespheginasister-group to [Sphegina + Neoascia] (Thompson, 1972, p.115). sister-group to Neoascia
In the chrysogasterines (Thompson et al., 1986). In the 
Spheginina subtribe (Thompson, 1989) 
Chamaesyrphussister-group to Pelecocera group, close to Rhingia sister-group to Pelecocera
(Thompson, 1972, p.102). In the Pelecocerina subtribe of the Rhingiini,
which also contains Cheilosia and Rhingia (Thompson, 1989)
 
ChasmommaAn eristaline (Hippa, 1978, p.14). Placed in the Helophilina in Helophilus group
subtribe (Thompson, 1989) 
Chromocheilosiaclose to the chrysogasterines (Thompson, 1972, p.115). in basal chrysogasterines
Related to Brachyopa and the chrysogasterines (Thompson, 1989) 
ChrysosyrphusRelated to Brachyopa and the chrysogasterines (Thompson, 1989)in the polytomy close to Brachyopa
Citrogrammastrong superficial resemblence to Xanthogramma [...] "but probably only real suggestion is as sister-group to
not closely related" (Vockeroth, 1969, p.94)Xanthogramma hence tentatively placed there
Cynorhinellain Myolepta group (Hippa, 1978, p.20). In Blera - Somula - sister-group to Lejota
Caliprobola - Lejota group (Thompson, 1989) 
Cyphipeltain the chrysogasterines (Thompson & Vockeroth, 1989). in the polytomy close to Brachyopa
Related to Brachyopa and the chrysogasterines (Thompson, 1989) 
Deinechessister-group to Flukea, and close to Criorhina sister-group to Flukea, with Malometasternum, close to Milesia
(Thompson, 1972, p.165). Sister-group to Flukea with 
Malometasternum; remarkable affinity to Milesia (Hippa, 1990, p.17) 
Dideoidessuperficially very similar to Didea (Vockeroth, 1969)in Didea group
Dideomimasister-group to Asiodidea (Vockeroth, 1969, p.106)in Didea group
Dideopsispossibly related to Asarkina (Vockeroth, 1969, p.113)in Didea group
Diguliarelated to Eristalis (Thompson & Vockeroth, 1989)in Eristalis group
Dissopterarelated to Eristalis (Thompson & Vockeroth, 1989)in Eristalis group
Dolichogynasimilar to Asemosyrphus and Arctosyrphus. in Helophilus group
(Thompson, 1972, p.134). Related to Helophilus 
(Thompson et al., 1976) 
Endoiasimyiarelated to Cheilosia - Ferdinandea - Rhingia (Knutson et al., 1975)in Cheilosia group
Eosphaerophoriain the Syrphini (Thompson & Vockeroth, 1989). placed in Baccha-Platycheirus-Sphaerophoria group
Among the most aberrant of all Syrphini; has been referred to at the boundary between the old Melanostomini and Syrphini
Melanostomini, but limits of that group are uncertain 
(Vockeroth, 1969, p.135) 
Eurimyiarelated to Anasimyia, in the Helophilina (Thompson, 1989)in Anasimyia group
Exallandrarelated to Melangyna (Vockeroth, 1969, p.87)sister-group to Melangyna
Flukeasister-group to Deineches, and close to Criorhina sister-group to Deineches (q.v.)
(Thompson, 1972, p.165). Sister-group to Deineches 
(Hippa, 1978, p.17) 
Giluweain the Syrphini, but very peculiar, without close relatives placed in Baccha-Platycheirus-Sphaerophoria group
[from description seems most like Eosphaerophoria] at the boundary between the old Melanostomini and Syrphini
(Vockeroth, 1969, p.136) 
Habromyiasister group to Mallota (Thompson, 1972, p.195). in Helophilus group
Related to Helophilus (Thompson et al., 1976). In the 
Helophilina (Thompson, 1989) 
Hadromyiasister-group to the Brachypalpoides group (Hippa, 1978), sister-group to the Brachypalpoides group
or to Sterphus (Thompson, 1972, p.195) 
Hemilamprain the chrysogasterines (Thompson & Vockeroth, 1989) with placed in the polytomy close to Brachyopa
Cyphipelta (Thompson, 1972 p.207) 
Hemixylotawith Stilbosoma, sister-group to Milesia (Thompson, 1972, p.194). in Milesia group
Relationship unknown (Hippa, 1978, p.18) 
Hiatomyiasister-group to Cheilosia sens.lat. (Thompson, 1972, p.102)in Cheilosia group
Ischyropterasister-group to [Pelecocera + Macropelecocera] in Pelecocera group
(Thompson, 1972, p.102) 
Kedarelated to Eristalis (Thompson & Vockeroth, 1989)in Eristalis group
Lepidomyiasister-group to Myolepta (Thompson, 1972, p.115)sister-group to Myolepta
Leucopodellain the Melanostomini (Thompson et al., 1976). Related to placed in Baccha-Platycheirus-Sphaerophoria group
Platycheirus-Baccha (Vockeroth, 1990)at the boundary between the old Melanostomini and Syrphini
LiochrysogasterRelated to Brachyopa and the chrysogasterines (Thompson, 1989)placed in basal chrysogasterines
Loveridgeanaa syrphine from St Helena (Smith & Vockeroth, 1980); placed in Baccha-Platycheirus-Sphaerophoria group
affinities uncertainat the boundary between the old Melanostomini and Syrphini
Lunomyiaplaced in same genus as Anasimyia (Thompson, 1989) in Anasimyia group
LycastrirhynchaRelated to Eristalis (Thompson, 1972 p.144, 195)in Eristalis group
Lycastrisclose to Criorhina (Hippa, 1978, p.19)in Criorhina group
MacrometopiaSubtribe Xylotina (Thompson, 1989). Related to Sterphus placed in polytomy close to Brachyopa
(Thompson, 1972, p.156); sister group of Xylota group 
(Thompson, 1972, p.194). NOT a xylotine, nor even close 
(Hippa 1978, p.19). 
Macropelecocerasister-group to Pelecocera (Thompson, 1972, p.102)in Pelecocera group
Macrozelimarelated to Spilomyia and Teuchocnemis (Hippa, 1978, p.19)with Teuchocnemis, in sister-group to Spilomyia
Malometasternumsister-group to [Deineches + Flukea] (Hippa, 1978, p.17)sister-group to [Deineches + Flukea]
Matsumyiaclosely related to Criorhina (Hippa, 1978, p.16)in Criorhina group
Melanogastersister-group to Chrysogaster (Maibach & Goeldlin, 1994)sister-group to Chrysogaster
Merapioidusallied to Criorhina group (Hippa, 1978, p.19)in Criorhina group
Meromacroidesrelated to Eristalis (Smith & Vockeroth, 1980)in Eristalis group
Meropidiain the Tropidiina subtribe (Thompson, 1989)sister-group to Tropidia
Mesembriusrelated to Helophilus (Thompson & Vockeroth, 1989)in Helophilus group
Mixogasterrelated to Microdon (Thompson et al., 1976)in Microdon group
NausigasterThompson (1972, p.125) and Hippa (1978, p.18) sister-group of Alipumilio
place it near Eumerus 
Nothomicrodonrelated to Microdon (Thompson et al., 1976)in Microdon group
Notiocheilosiaprobably related to Callicera (Thompson, 1972, p.113, 202)sister-group to Callicera
Notosyrphussister group of Dasysyrphus (Vockeroth, 1969, p.73, 75)sister-group to Dasysyrphus
Odyneromyiasister-group to Valdiviomyia (Thompson, 1972, p.154) and sister-group to Valdiviomyia, then to [Temnostoma + Takaomyia]
related to Temnostoma group (Hippa, 1978, p.22) 
OrthoprosopaIn the Criorhina group (Thompson, 1972). In in Criorhina group
Pterallastina xylotines (Thompson & Vockeroth, 1989) 
Palpadaformerly in Eristalis [hence closely related] (Thompson, 1972, p.139)in Eristalis group
Palumbiasister-group to Pterallastes, and related to Syrittosyrphus and sister-group to Pterallastes
Milesia (Hippa, 1978, p.24) 
Paragodonrelated to Microdon (Thompson et al., 1976)in Microdon group
Paramesembriusrelated to Helophilus (Thompson, 1989)in Helophilus group
Paramicrodonrelated to Microdon (Thompson et al., 1976)in Microdon group
Paramixogasterrelated to Microdon (Thompson, 1989)in Microdon group
Paramixogasteroidesrelated to Microdon (Thompson, 1989)in Microdon group
Pararctophilain the Sericomyiini (Thompson, 1972, 206)in Sericomyia group
Pelecoceraclose to Rhingia (Thompson, 1972, p.102). In the in Pelecocera group
Pelecocerina subtribe of the Rhingiini, which also contains 
Cheilosia and Rhingia (Thompson, 1989) 
Pellolomain the Syrphini (Smith & Vockeroth, 1980)placed in Baccha-Platycheirus-Sphaerophoria group at the boundary between the old Melanostomini and Syrphini
Philippimyiaa xylotine, perhaps related to Stilbosoma (Thompson, 1972, p.166); sister-group to Blera
sister group of Blera (Thompson, 1972, p.194). 
Apparently related to Blera (Hippa, 1978, p.23) 
Phytomiarelated to Eristalis (Thompson & Vockeroth, 1989)in Eristalis group
Piaunknown affinities (Thompson, 1989)omitted
Pilinasciasister-group to Helophilus (Thompson & Vockeroth, 1989)sister-group to Helophilus
Pogonosyrphussister-group to Milesia (Hippa, 1978, p.20)sister-group to Milesia
Polydontomyiarelated to Helophilus (Thompson et al., 1976). Placed in the in Anasimyia group
same genus as Anasimyia (Thompson, 1989) 
Primocerioidesrelated to Ceriana (Thompson, 1989)sister-group to Ceriana
Psarocheilosiarelated to Pelecocera (Thompson, 1989)in Pelecocera group
Psarusin its own tribe (Thompson, 1972, p.202). Related to in Cheilosia group
Cheilosia-Rhingia (Thompson, 1989) 
Pseuderistalisrelated to Eristalis (Thompson & Vockeroth, 1989)in Eristalis group
Pseudovolucellain the Sericomyiini (Thompson, 1972, 202)in Sericomyia group
Pterallastessister-group to Palumbia, related to Syrittosyrphus sister-group to Palumbia
(Hippa, 1978, p.24) 
Pyritisin the Sericomyiini (Thompson, 1972, 202)in Sericomyia group
Rhinobaccharelated to Allobaccha (Knutson et al., 1975)placed in Baccha-Platycheirus-Sphaerophoria group at the boundary between the old Melanostomini and Syrphini
Rhysopsin the Melanostomini (Thompson et al., 1976)placed in Baccha-Platycheirus-Sphaerophoria group at the boundary between the old Melanostomini and Syrphini
Senaspisrelated to Eristalis (Smith & Vockeroth, 1980)in Eristalis group
Senogastersister group of Syritta (Thompson, 1972, p.169). In sister-group to Nepenthosyrphus, and then to Syritta
Tropidia group (Hippa, 1978, p.28) 
Simoidesrelated to Eristalis (Smith & Vockeroth, 1980)in Eristalis group
Simosyrphussister-group to Ischiodon (Vockeroth, 1969, p.105). sister-group to Ischiodon
In Eupeodes group (Dusek & Laska, 1985) 
Solenaspisrelated to Eristalis (Thompson & Vockeroth, 1989)in Eristalis group
Spazigasterrelated to Melanostoma-Xanthandrus (Dusek & Laska, 1967)placed in Baccha-Platycheirus-Sphaerophoria group at the boundary between the old Melanostomini and Syrphini
SphecomyiaClosely related to Criorhina (Hippa, 1978, p.16)grouped with [ Criorhina + Matsumyia ]
Spheginobacchain its own tribe (Thompson, 1972, p.202). Similar features to placed in polytomy close to Brachyopa
Neoascia, Sphegina, Syritta and Baccha (Brunetti, 1915) 
Stenopipizaunknown affinities (Thompson, 1989)omitted
Sterphusrelated to Xylota group and Macrometopia sister-group to Xylota
(Thompson, 1972, p.156); sister group to Hadromyia 
(Thompson, 1972, p.195). Sister group to Xylota (Hippa, 1978) 
Stilbosomaa xylotine (Thompson, 1972, p.174). Related to Milesia in Milesia group
(Hippa, 1978, p.25) 
Syrittosyrphussister -group to [ Palumbia + Pterallastes ] (Hippa, 1978, p.24)in Milesia group
Syrphocheilosiain Melanostomini (Thompson, 1989)placed in Baccha-Platycheirus-Sphaerophoria group at the boundary between the old Melanostomini and Syrphini
Taeniocheilosiarelated to Pelecocera (Thompson, 1989)in Pelecocera group
Takaomyiabelonging to the Temnostoma group (Thompson, 1972, p.154). sister-group to Temnostoma
Sister-group to Temnostoma (Hippa, 1978, p.25) 
Talahuain the Melanostomini (Thompson et al., 1976)placed in Baccha-Platycheirus-Sphaerophoria group at the boundary between the old Melanostomini and Syrphini
Teuchocnemispossibly related to Aneriophora (Hippa, 1978, p.11); in Spilomyia group
related to Spilomyia and Macrozelima (Hippa, 1978, p.19) 
Tuberculanostomain the Melanostomini (Thompson et al., 1976)placed in Baccha-Platycheirus-Sphaerophoria group at the boundary between the old Melanostomini and Syrphini
Valdiviomyiasister-group of Odyneromyia, related to Temnostoma
(Thompson, 1972, p.154; Hippa, 1978, p.22)
in Temnostoma group, sister-group of Odyneromyia
#NEXUS
[MacClade 3.05 registered to pharv, zoology]
BEGIN TAXA;
DIMENSIONS NTAX=208;
TAXLABELS
_35_Lonchopteridae
215_Platypezidae
2500_Phoridae
600_Pipunculidae
258_Eumerus
_15_Nausigaster
__4_Alipumilio
__1_Austrocheilosia
104_Merodon
__7_Azypetia
__6_Chamaesyrphus
__6_Pelecocera
__1_Ischyroptera
__1_Macropelecocera
__1_Psarocheilosia
__1_Taeniocheilosia
__?_Cheilosia_miners
__3_Portevinia
390_Cheilosia_borers
__?_Cheilosia_paganus
_21_Hiatomyia
__6_Endoiasimyia
__1_Psarus
__?_Cheilosia_morio
__?_Cheilosia_fungivores
_12_Ferdinandea
_37_Rhingia
_77_Graptomyza
325_Copestylum
__3_Ornidia
__1_Volucella_inflata
__5_Volucella_inanis
_18_Volucella_pellucens
_18_Volucella_bombylans
358_Microdon
_20_Mixogaster
_24_other_Microdons
_27_Pipizella
_34_Neocnemodon
_25_Trichopsomyia
_47_Pipiza
_10_Heringia
_52_Melanostoma
_25_Xanthandrus
148_Toxomerus
141_Platycheirus
300_Ocyptamus
_16_Leucopodella
_31_Rhysops
__2_Spazigaster
__1_Syrphocheilosia
__1_Talahua
__5_Tuberculanostoma
__4_Pyrophaena
__5_Rohdendorfia
__2_Eosphaerophoria
__2_Giluwea
__1_Loveridgeana
__1_Pelloloma
__1_Pseudodoros
__1_Rhinobaccha
_35_Salpingogaster
_23_Baccha
_58_Sphaerophoria
114_Allograpta
_19_Meliscaeva
_26_Episyrphus
_16_Betasyrphus
_21_Parasyrphus
__1_Fagisyrphus
_22_Syrphus
_31_Melangyna
__1_Exallandra
_90_Chrysotoxum
_61_Paragus
_71_Eupeodes
_16_Scaeva
__4_Ischiodon
__1_Simosyrphus
_36_Dasysyrphus
__1_Notosyrphus
__3_Eriozona
__4_Didea
_44_Asarkina
__1_Asiodidea
_11_Dideoides
__1_Dideomima
__3_Dideopsis
__4_Megasyrphus
__5_Meligramma
__3_Epistrophella
_20_Xanthogramma
_17_Citrogramma
__4_Doros
_21_Epistrophe
__1_Afrosyrphus
__9_Leucozona
__2_Hammerschmidtia
_29_Brachyopa
__3_Cacoceria
__6_Chrysosyrphus
__1_Cyphipelta
__4_Hemilampra
_14_Spheginobaccha
__2_Macrometopia
__2_Chromocheilosia
__1_Liochrysogaster
__4_Lejogaster
_23_Chrysogaster
_12_Melanogaster
_55_Orthonevra
__2_Chamaesphegina
__1_Austroascia
_19_Neoascia
_68_Sphegina
_95_Xylota
_37_Sterphus
_24_Tropidia
__3_Meropidia
_26_Syritta
__1_Senogaster
__5_Nepenthosyrphus
_32_Psilota
187_Ceriana
__1_Primocerioides
__3_Pocota
_28_Blera
__1_Philippimyia
__2_Caliprobola
__2_Somula
__8_Lejota
__2_Cynorhinella
_34_Myolepta
_16_Lepidomyia
_28_Sericomyia
__1_Pararctophila
__8_Pseudovolucella
__1_Pyritis
_56_Mallota
_98_Eristalinus
_23_Austalis
__2_Axona
__1_Digulia
__7_Dissoptera
__1_Keda
__1_Meromacroides
_95_Palpada
_43_Meromacrus
__5_Lycastrirhyncha
_24_Phytomia
_17_Pseuderistalis
_14_Senaspis
__7_Simoides
__1_Solenaspis
__2_Eristalis
_81_Eoseristalis
__3_Myathropa
_11_Dolichogyna
_12_Pilinascia
_25_Helophilus
_47_Mesembrius
__1_Paramesembrius
_25_Quichuana
_11_Habromyia
__3_Austrophilus
__3_Chasmomma
_19_Anasimyia
__1_Arctosyrphus
__3_Asemosyrphus
__2_Eurimyia
__2_Lejops
__1_Lunomyia
__1_Polydontomyia
_17_Parhelophilus
_57_Chalcosyrphus
_41_Xylotomima
__1_Ameroxylota
_12_Brachypalpus
_20_Brachypalpoides
__7_Hadromyia
_31_Spilomyia
__1_Macrozelima
_2_Teuchocnemis
__1_Aneriophora
__4_Pterallastes
_17_Palumbia
_49_Milesia
__3_Pogonosyrphus
__2_Stilbosoma
__3_Hemixylota
__1_Syrittosyrphus
__1_Malometasternum
__1_Flukea
__3_Deineches
_51_Criorhina
__8_Matsumyia
__9_Sphecomyia
__6_Orthoprosopa
__1_Merapioidus
__5_Lycastris
__6_Odyneromyia
__6_Valdiviomyia
_17_Temnostoma
__3_Takaomyia
_18_Callicera
__1_Notiocheilosia
__5_Triglyphus
_75_Allobaccha;
END;
BEGIN CHARACTERS;
DIMENSIONS NCHAR=2;
FORMAT SYMBOLS= " 0 1 2 3 4 5" MISSING=? GAP=- ;OPTIONS MSTAXA=POLYMORPH ;
MATRIX
_35_Lonchopteridae  ??
215_Platypezidae  ??
2500_Phoridae  ??
600_Pipunculidae  ??
258_Eumerus  ?(1235)
_15_Nausigaster  ?(04)
__4_Alipumilio  ?0
__1_Austrocheilosia  ?5
104_Merodon  ?(123)
__7_Azypetia  ?(135)
__6_Chamaesyrphus  ??
__6_Pelecocera  ?(014)
__1_Ischyroptera  ?1
__1_Macropelecocera  ?1
__1_Psarocheilosia  ?1
__1_Taeniocheilosia  ?1
__?_Cheilosia_miners  ??
__3_Portevinia  ?1
390_Cheilosia_borers  ??
__?_Cheilosia_paganus  ??
_21_Hiatomyia  ?4
__6_Endoiasimyia  ?(13)
__1_Psarus  ?1
__?_Cheilosia_morio  ?1
__?_Cheilosia_fungivores  ??
_12_Ferdinandea  ?(134)
_37_Rhingia  ?(01234)
_77_Graptomyza  ?(1235)
325_Copestylum  ?(04)
__3_Ornidia  ?0
__1_Volucella_inflata  ?1
__5_Volucella_inanis  ?1
_18_Volucella_pellucens  ?(13)
_18_Volucella_bombylans  ?(134)
358_Microdon  ?(012345)
_20_Mixogaster  ?(04)
_24_other_Microdons  ?(023)
_27_Pipizella  ?1
_34_Neocnemodon  ?(14)
_25_Trichopsomyia  ?(014)
_47_Pipiza  ?(014)
_10_Heringia  ?(134)
_52_Melanostoma  ?(012345)
_25_Xanthandrus  ?(012345)
148_Toxomerus  ?(04)
141_Platycheirus  ?(01345)
300_Ocyptamus  ?(04)
_16_Leucopodella  ?(04)
_31_Rhysops  ?0
__2_Spazigaster  ?1
__1_Syrphocheilosia  ?1
__1_Talahua  ?0
__5_Tuberculanostoma  ?(01)
__4_Pyrophaena  ??
__5_Rohdendorfia  ?1
__2_Eosphaerophoria  ?3
__2_Giluwea  ?5
__1_Loveridgeana  ?2
__1_Pelloloma  ?2
__1_Pseudodoros  ?(04)
__1_Rhinobaccha  ?3
_35_Salpingogaster  ?(04)
_23_Baccha  ?(1234)
_58_Sphaerophoria  ?(12345)
114_Allograpta  ?(012345)
_19_Meliscaeva  ?(134)
_26_Episyrphus  ?(1235)
_16_Betasyrphus  ?(1235)
_21_Parasyrphus  ?(134)
__1_Fagisyrphus  ?1
_22_Syrphus  ?(012345)
_31_Melangyna  ?(1345)
__1_Exallandra  ?2
_90_Chrysotoxum  ?(01234)
_61_Paragus  ?(012345)
_71_Eupeodes  ?(012345)
_16_Scaeva  ?(0134)
__4_Ischiodon  ?(1235)
__1_Simosyrphus  ?5
_36_Dasysyrphus  ?(134)
__1_Notosyrphus  ?0
__3_Eriozona  ?(134)
__4_Didea  ?(14)
_44_Asarkina  ?(1235)
__1_Asiodidea  ?1
_11_Dideoides  ?(13)
__1_Dideomima  ?(04)
__3_Dideopsis  ?(35)
__4_Megasyrphus  ?(14)
__5_Meligramma  ?(14)
__3_Epistrophella  ?(145)
_20_Xanthogramma  ?(134)
_17_Citrogramma  ?(35)
__4_Doros  ?(14)
_21_Epistrophe  ?(134)
__1_Afrosyrphus  ?2
__9_Leucozona  ?(134)
__2_Hammerschmidtia  ?(14)
_29_Brachyopa  ?(14)
__3_Cacoceria  ?0
__6_Chrysosyrphus  ?(014)
__1_Cyphipelta  ?5
__4_Hemilampra  ?5
_14_Spheginobaccha  ?(23)
__2_Macrometopia  ?0
__2_Chromocheilosia  ?0
__1_Liochrysogaster  ?1
__4_Lejogaster  ?1
_23_Chrysogaster  ?(1234)
_12_Melanogaster  ?1
_55_Orthonevra  ?(0134)
__2_Chamaesphegina  ?0
__1_Austroascia  ?0
_19_Neoascia  ?(14)
_68_Sphegina  ?(134)
_95_Xylota  ?(012345)
_37_Sterphus  ?0
_24_Tropidia  ?(0124)
__3_Meropidia  ?0
_26_Syritta  ?(012345)
__1_Senogaster  ?0
__5_Nepenthosyrphus  ?3
_32_Psilota  ?(1345)
187_Ceriana  ?(012345)
__1_Primocerioides  ??
__3_Pocota  ?(14)
_28_Blera  ?(14)
__1_Philippimyia  ?0
__2_Caliprobola  ?1
__2_Somula  ?4
__8_Lejota  ?(14)
__2_Cynorhinella  ?4
_34_Myolepta  ?(01234)
_16_Lepidomyia  ?0
_28_Sericomyia  ?(0134)
__1_Pararctophila  ?1
__8_Pseudovolucella  ?(13)
__1_Pyritis  ?4
_56_Mallota  ?(01234)
_98_Eristalinus  ?(012345)
_23_Austalis  ?(35)
__2_Axona  ?(35)
__1_Digulia  ?5
__7_Dissoptera  ?(35)
__1_Keda  ?(35)
__1_Meromacroides  ?2
_95_Palpada  ?(04)
_43_Meromacrus  ?(04)
__5_Lycastrirhyncha  ?0
_24_Phytomia  ?(1235)
_17_Pseuderistalis  ?(135)
_14_Senaspis  ?2
__7_Simoides  ?2
__1_Solenaspis  ?5
__2_Eristalis  ?(012345)
_81_Eoseristalis  ??
__3_Myathropa  ?1
_11_Dolichogyna  ?0
_12_Pilinascia  ?5
_25_Helophilus  ?(01345)
_47_Mesembrius  ?(1235)
__1_Paramesembrius  ?(13)
_25_Quichuana  ?0
_11_Habromyia  ?0
__3_Austrophilus  ?5
__3_Chasmomma  ?2
_19_Anasimyia  ?(124)
__1_Arctosyrphus  ?(14)
__3_Asemosyrphus  ?(04)
__2_Eurimyia  ?(14)
__2_Lejops  ?(14)
__1_Lunomyia  ?4
__1_Polydontomyia  ?(04)
_17_Parhelophilus  ?(14)
_57_Chalcosyrphus  ?(01345)
_41_Xylotomima  ?(14)
__1_Ameroxylota  ?4
_12_Brachypalpus  ?(14)
_20_Brachypalpoides  ?(13)
__7_Hadromyia  ?(14)
_31_Spilomyia  ?(0134)
__1_Macrozelima  ?1
_2_Teuchocnemis  ?4
__1_Aneriophora  ?0
__4_Pterallastes  ?(14)
_17_Palumbia  ?(134)
_49_Milesia  ?(01234)
__3_Pogonosyrphus  ?2
__2_Stilbosoma  ?0
__3_Hemixylota  ?0
__1_Syrittosyrphus  ?2
__1_Malometasternum  ?5
__1_Flukea  ?0
__3_Deineches  ?5
_51_Criorhina  ?(0134)
__8_Matsumyia  ?(13)
__9_Sphecomyia  ?(14)
__6_Orthoprosopa  ?5
__1_Merapioidus  ?4
__5_Lycastris  ?3
__6_Odyneromyia  ?(05)
__6_Valdiviomyia  ?0
_17_Temnostoma  ?(134)
__3_Takaomyia  ?(13)
_18_Callicera  ?(0134)
__1_Notiocheilosia  ?0
__5_Triglyphus  ?(135)
_75_Allobaccha  ?(1235)
;
END;
BEGIN ASSUMPTIONS;
OPTIONS DEFTYPE=unord PolyTcount=MINSTEPS ;
END;
BEGIN TREES;
TRANSLATE
  1  _35_Lonchopteridae,
  2  215_Platypezidae,
  3  2500_Phoridae,
  4  600_Pipunculidae,
  5  258_Eumerus,
  6  _15_Nausigaster,
  7  __4_Alipumilio,
  8  __1_Austrocheilosia,
  9  104_Merodon,
  10  __7_Azypetia,
  11  __6_Chamaesyrphus,
  12  __6_Pelecocera,
  13  __1_Ischyroptera,
  14  __1_Macropelecocera,
  15  __1_Psarocheilosia,
  16  __1_Taeniocheilosia,
  17  __?_Cheilosia_miners,
  18  __3_Portevinia,
  19  390_Cheilosia_borers,
  20  __?_Cheilosia_paganus,
  21  _21_Hiatomyia,
  22  __6_Endoiasimyia,
  23  __1_Psarus,
  24  __?_Cheilosia_morio,
  25  __?_Cheilosia_fungivores,
  26  _12_Ferdinandea,
  27  _37_Rhingia,
  28  _77_Graptomyza,
  29  325_Copestylum,
  30  __3_Ornidia,
  31  __1_Volucella_inflata,
  32  __5_Volucella_inanis,
  33  _18_Volucella_pellucens,
  34  _18_Volucella_bombylans,
  35  358_Microdon,
  36  _20_Mixogaster,
  37  _24_other_Microdons,
  38  _27_Pipizella,
  39  _34_Neocnemodon,
  40  _25_Trichopsomyia,
  41  _47_Pipiza,
  42  _10_Heringia,
  43  _52_Melanostoma,
  44  _25_Xanthandrus,
  45  148_Toxomerus,
  46  141_Platycheirus,
  47  300_Ocyptamus,
  48  _16_Leucopodella,
  49  _31_Rhysops,
  50  __2_Spazigaster,
  51  __1_Syrphocheilosia,
  52  __1_Talahua,
  53  __5_Tuberculanostoma,
  54  __4_Pyrophaena,
  55  __5_Rohdendorfia,
  56  __2_Eosphaerophoria,
  57  __2_Giluwea,
  58  __1_Loveridgeana,
  59  __1_Pelloloma,
  60  __1_Pseudodoros,
  61  __1_Rhinobaccha,
  62  _35_Salpingogaster,
  63  _23_Baccha,
  64  _58_Sphaerophoria,
  65  114_Allograpta,
  66  _19_Meliscaeva,
  67  _26_Episyrphus,
  68  _16_Betasyrphus,
  69  _21_Parasyrphus,
  70  __1_Fagisyrphus,
  71  _22_Syrphus,
  72  _31_Melangyna,
  73  __1_Exallandra,
  74  _90_Chrysotoxum,
  75  _61_Paragus,
  76  _71_Eupeodes,
  77  _16_Scaeva,
  78  __4_Ischiodon,
  79  __1_Simosyrphus,
  80  _36_Dasysyrphus,
  81  __1_Notosyrphus,
  82  __3_Eriozona,
  83  __4_Didea,
  84  _44_Asarkina,
  85  __1_Asiodidea,
  86  _11_Dideoides,
  87  __1_Dideomima,
  88  __3_Dideopsis,
  89  __4_Megasyrphus,
  90  __5_Meligramma,
  91  __3_Epistrophella,
  92  _20_Xanthogramma,
  93  _17_Citrogramma,
  94  __4_Doros,
  95  _21_Epistrophe,
  96  __1_Afrosyrphus,
  97  __9_Leucozona,
  98  __2_Hammerschmidtia,
  99  _29_Brachyopa,
  100  __3_Cacoceria,
  101  __6_Chrysosyrphus,
  102  __1_Cyphipelta,
  103  __4_Hemilampra,
  104  _14_Spheginobaccha,
  105  __2_Macrometopia,
  106  __2_Chromocheilosia,
  107  __1_Liochrysogaster,
  108  __4_Lejogaster,
  109  _23_Chrysogaster,
  110  _12_Melanogaster,
  111  _55_Orthonevra,
  112  __2_Chamaesphegina,
  113  __1_Austroascia,
  114  _19_Neoascia,
  115  _68_Sphegina,
  116  _95_Xylota,
  117  _37_Sterphus,
  118  _24_Tropidia,
  119  __3_Meropidia,
  120  _26_Syritta,
  121  __1_Senogaster,
  122  __5_Nepenthosyrphus,
  123  _32_Psilota,
  124  187_Ceriana,
  125  __1_Primocerioides,
  126  __3_Pocota,
  127  _28_Blera,
  128  __1_Philippimyia,
  129  __2_Caliprobola,
  130  __2_Somula,
  131  __8_Lejota,
  132  __2_Cynorhinella,
  133  _34_Myolepta,
  134  _16_Lepidomyia,
  135  _28_Sericomyia,
  136  __1_Pararctophila,
  137  __8_Pseudovolucella,
  138  __1_Pyritis,
  139  _56_Mallota,
  140  _98_Eristalinus,
  141  _23_Austalis,
  142  __2_Axona,
  143  __1_Digulia,
  144  __7_Dissoptera,
  145  __1_Keda,
  146  __1_Meromacroides,
  147  _95_Palpada,
  148  _43_Meromacrus,
  149  __5_Lycastrirhyncha,
  150  _24_Phytomia,
  151  _17_Pseuderistalis,
  152  _14_Senaspis,
  153  __7_Simoides,
  154  __1_Solenaspis,
  155  __2_Eristalis,
  156  _81_Eoseristalis,
  157  __3_Myathropa,
  158  _11_Dolichogyna,
  159  _12_Pilinascia,
  160  _25_Helophilus,
  161  _47_Mesembrius,
  162  __1_Paramesembrius,
  163  _25_Quichuana,
  164  _11_Habromyia,
  165  __3_Austrophilus,
  166  __3_Chasmomma,
  167  _19_Anasimyia,
  168  __1_Arctosyrphus,
  169  __3_Asemosyrphus,
  170  __2_Eurimyia,
  171  __2_Lejops,
  172  __1_Lunomyia,
  173  __1_Polydontomyia,
  174  _17_Parhelophilus,
  175  _57_Chalcosyrphus,
  176  _41_Xylotomima,
  177  __1_Ameroxylota,
  178  _12_Brachypalpus,
  179  _20_Brachypalpoides,
  180  __7_Hadromyia,
  181  _31_Spilomyia,
  182  __1_Macrozelima,
  183  _2_Teuchocnemis,
  184  __1_Aneriophora,
  185  __4_Pterallastes,
  186  _17_Palumbia,
  187  _49_Milesia,
  188  __3_Pogonosyrphus,
  189  __2_Stilbosoma,
  190  __3_Hemixylota,
  191  __1_Syrittosyrphus,
  192  __1_Malometasternum,
  193  __1_Flukea,
  194  __3_Deineches,
  195  _51_Criorhina,
  196  __8_Matsumyia,
  197  __9_Sphecomyia,
  198  __6_Orthoprosopa,
  199  __1_Merapioidus,
  200  __5_Lycastris,
  201  __6_Odyneromyia,
  202  __6_Valdiviomyia,
  203  _17_Temnostoma,
  204  __3_Takaomyia,
  205  _18_Callicera,
  206  __1_Notiocheilosia,
  207  __5_Triglyphus,
  208  _75_Allobaccha
;
TREE T1 = [&R]
(1,(2,(3,(4,((5,8,(6,7)),(9,10),(13,14,15,16,(11,12)),(17,((18,(19,20,21,22,
23)),(24,(25,((26,27),((28,29,30,31),(((32,(33,34)),((35,36,37),((38,(39,40,
(41,42))),(43,(44,(45,46,47,48,49,50,51,52,53,56,57,58,59,60,61,62,(54,55),(
63,((64,65),((66,(67,68)),(69,(70,((71,(72,73)),((74,((75,(76,(77,(78,79))))
,((80,81),(82,(89,(83,84,85,86,87,88)))))),(((90,91),(94,(92,93))),(97,(95,9
6)))))))))))))))),(100,101,102,103,104,105,(98,99),((106,107,(108,(111,(109,
110)))),((112,113,(114,115)),((116,117),((118,119,(123,(120,(121,122)))),((1
24,125),((126,((129,130,(127,128)),((131,132),((133,134),((135,136,137,138),
(139,(140,((141,142,143,144,145,146,147,148,149,150,151,152,153,154,(157,(15
5,156))),(158,161,162,163,164,165,166,(159,160),(174,(167,168,169,170,171,17
2,173))))))))))),(((175,176),(180,(179,(177,178)))),(((181,(182,183,184)),((
189,190,191,((185,186),(187,188))),(192,(193,194)))),((198,199,200,(195,196,
197)),(((201,202),(203,204)),(205,206))))))))))))))))))))))));
TREE * UNTITLED = [&R]
(1,(2,(3,(4,((5,8,(6,7)),(9,10),(13,14,15,16,(11,12)),(17,((18,(19,20,21,22,
23)),(24,(25,((26,27),((28,29,30,31),(((32,(33,34)),((35,36,37),(((38,207),(
39,40,(41,42))),(43,(44,(45,46,47,48,49,50,51,52,53,56,57,58,59,60,61,62,(54
,55),((63,208),((64,65),((66,(67,68)),(69,(70,((71,(72,73)),((74,((75,(76,(7
7,(78,79)))),((80,81),(82,(89,(83,84,85,86,87,88)))))),(((90,91),(94,(92,93)
)),(97,(95,96)))))))))))))))),(100,101,102,103,104,105,(98,99),((106,107,(10
8,(111,(109,110)))),((112,113,(114,115)),((116,117),((118,119,(123,(120,(121
,122)))),((124,125),((126,((129,130,(127,128)),((131,132),((133,134),((135,1
36,137,138),(139,(140,((141,142,143,144,145,146,147,148,149,150,151,152,153,
154,(157,(155,156))),(158,161,162,163,164,165,166,(159,160),(174,(167,168,16
9,170,171,172,173))))))))))),(((175,176),(180,(179,(177,178)))),(((181,(182,
183,184)),((189,190,191,((185,186),(187,188))),(192,(193,194)))),((198,199,2
00,(195,196,197)),(((201,202),(203,204)),(205,206))))))))))))))))))))))));
END;
BEGIN MACCLADE;
Version 3.05;
LastModified -1298882056;
Singles 0100&/0;
END;